- this may have been a battle between two Central European armies, one rich in Y-haplogroup R1b and the other rich in Y-haplogroup I2a, as well as their allies or hired help, including warriors from Eastern Europe belonging to Y-haplogroup R1a - or perhaps it was an invasion from the east by warriors rich in Y-haplogroup R1a, and it was a success, with the local armies, rich in Y-haplogroups R1b and I2a, losing the battle and suffering most of the casualties.I'm sure that one day someone will attempt to undertake a decent multidisciplinary study of this epic battle, and we'll at least have a rough idea about what happened. Or not. Citation... Burger et al., Low Prevalence of Lactase Persistence in Bronze Age Europe Indicates Ongoing Strong Selection over the Last 3,000 Years, Current Biology, Available online 3 September 2020, https://doi.org/10.1016/j.cub.2020.08.033 See also... Genetic and linguistic structure across space and time in Northern Europe
search this blog
Showing posts with label Bronze Age. Show all posts
Showing posts with label Bronze Age. Show all posts
Tuesday, September 8, 2020
Warriors from at least two different populations fought in the Tollense Valley battle
I can't get the genotype data from the Burger et al. paper. The lead authors, Joachim Burger and Daniel Wegmann, aren't replying to my emails.
But they were gracious enough to release the BAM files for each of their samples, and these files can be converted to genotype data. So I've included ten of the Tollense Valley warriors (DEU_Tollense_BA) in the Global25 datasheets (see here).
The claim in the paper that these warriors "represent an unstructured population" is absolutely false and extremely naive.
Below are a couple of Principal Component Analysis (PCA) plots produced with Vahaduo Global25 views. The samples are labeled according to their Y-chromosome haplogroups. To see interactive versions of the same plots, paste the Global25 coordinates from the text file here into the relevant fields here.
These warriors are not a single unstructured population, because they cover too much ground in the above plots for that to be possible. It's clear to me that they represent at least two different groups from Central Europe and surrounds.
Of course, this would be a lot easier to work out if Burger et al. cared to supply more information about each of the warriors, such as their attire, weapons, circumstances of death, and so on. It's a complete mystery to me why this wasn't included in the paper, and the authors are refusing to talk to me, so it's unlikely that I'll ever be able to get it from them.
In the absence of such crucial archeological and anthropological data, I don't want to speculate too much, and get overly creative, but here are a couple of possible scenarios to explain the ancient DNA results:
Labels:
ancient DNA,
Balto-Slavic,
battle,
Bronze Age,
Central Europe,
dead,
Eastern Europe,
Germany,
I2a,
Northern Europe,
Poland,
R1a,
R1a-Z283,
R1b,
R1b-P312,
Tollense Valley,
victims,
warlike
Sunday, March 31, 2019
Map of pre-Corded Ware culture (>2900 BCE) instances of Y-haplogroup R1a (updated)
Below is a map showing the global distribution of Y-chromosome haplogroup R1a prior to the expansions of the R1a-rich Corded Ware culture (CWC) people and their descendants across Europe and Asia from around 2900 BCE. I'll be updating this map regularly and using it to help me narrow down the options for the place of origin of R1a, and also to counter the misinformation about this topic that has appeared in print and online over the years, including in many scientific publications and popular websites such as Wikipedia.
Incredibly, as far as I know, there are just six reliably called instances of R1a in the now ample Eurasian ancient DNA record dating to the pre-CWC period. To put this into perspective, consider that R1a is today the most common Y-haplogroup in much of Europe and Asia. How did that happen I wonder? However, please note that I chose to base the map only on samples sequenced with the capture and shotgun methods, rather than the PCR method, which is susceptible to producing contaminated results and no longer used in major ancient DNA studies.
See also...
Y-haplogroup R1a and mental health
The Poltavka outlier
Late PIE ground zero now obvious; location of PIE homeland still uncertain, but...
Labels:
ancient DNA,
Bronze Age,
Copper Age,
Corded Ware Culture,
CWC,
Eastern Europe,
Eurasia,
PIE,
Proto-Indo-European,
R1a,
R1a origin,
R1a-M17,
R1a-M198,
R1a-M417,
R1a-M420,
R1a-Z645,
R1a-Z93,
R1a1a1,
Y-haplogroup
Monday, March 4, 2019
An exceptional burial indeed, but not that of an Indo-European
Not too many people have been buried sitting on wagons. The most famous case is that of an Early Bronze Age man who, considering his injuries, may have died in a high-speed crash - high-speed for its time anyway - on the Pontic-Caspian steppe in Eastern Europe.
It's likely that this guy was one of the very first wagon-drivers in human history, because his four-wheeled wooden model is dated to 3336-3105 calBCE, which makes it the oldest wagon discovered thus far. His genotype data, under the label Steppe Maykop SA6004, were published recently along with Wang et al. 2019.
Early wagons are very important for a couple of reasons: they revolutionized human transport and warfare, and they're often closely associated with the prehistoric expansions of Indo-European languages.
So I'm pretty sure that many of you must be thinking right now that wagon-driver SA6004 was an early Indo-European, or even a Proto-Indo-European! I bet that's what Wang et al. thought too, considering the conclusion in their paper. But, alas, the chances of this are slim to none.
Steppe Maykop samples show rather peculiar genetic structure considering their geographic origin, with a large proportion of their ancestry deriving from a source closely related to western Siberian hunter-gatherers (aka West_Siberia_N in the ancient DNA record). Indeed, SA6004 basically looks like a 50/50 mix between West_Siberia_N and Piedmont_Eneolithic. Here's a map with all of the relevant details.
Thus, clearly, the Steppe Maykop population wasn't ancestral or even directly related to the steppe and steppe-derived groups generally regarded to have been Indo-European speaking, such as those associated with the Yamnaya, Corded Ware, and Bell Beaker cultures. That's because these groups lack any discernible West_Siberia_N-related ancestry.
It also wasn't ancestral or directly related to any present-day or currently sampled ancient Indo-European speaking populations, again because these populations basically lack West_Siberia_N-related ancestry.
On the other hand, Yamnaya, Corded Ware and other closely related groups show an exceptionally strong genetic relationship with Indo-European speakers, especially those from across Northern Europe, which experienced massive migrations from the Pontic-Caspian steppe during the late Neolithic period, and hardly anything from elsewhere since then.
Case in point, the samples from Wang et al. labeled Yamnaya Caucasus were recovered from the same area of the Pontic-Caspian as their Steppe Maykop samples, and yet, take a look at this linear model based on outgroup f3-statistics. Steppe Maykop does show high genetic affinity to Indo-European speakers (no doubt mediated via its Piedmont_Eneolithic-related ancestry), but, unlike Yamnaya Caucasus, it also shows unusually high affinity for a West Eurasian population to Native Americans and Siberians. The relevant datasheet is available here.
So the only way that the Steppe Maykop population was Indo-European-speaking, was if it inherited its Indo-European speech from its Piedmont_Eneolithic-related ancestors. And even if it was Indo-European-speaking, it probably spoke an extinct Indo-European language not closely related to any extant Indo-European languages. In other words, the possibility that Steppe Maykop passed on its language to Yamnaya, along with its wagons, is close to zero. More likely, Yamnaya stole a few wagons from Steppe Maykop, and the rest is history.
See also...
The Steppe Maykop enigma
On Maykop ancestry in Yamnaya
Late PIE ground zero now obvious; location of PIE homeland still uncertain, but...
Labels:
ancient DNA,
Bronze Age,
Caucasus,
Indo-European,
kurgan,
Maykop,
migration,
PIE,
Pontic-Caspian steppe,
Progress Eneolithic,
qpAdm,
Siberia,
Steppe Maykop,
wagon,
wagon burial,
wheel,
Yamnaya
Saturday, March 2, 2019
Maykop: a multi-ethnic layer cake?
Let's speculate about the linguistic affinities of the currently available ancient populations from the Caucasus and surrounds. I put together a series of outgroup f3-stats to help things along. They're available for download here.
Maykop Georgian 0.258224 Abkhasian 0.257899 Latvian 0.257376 Swedish 0.257301 Turkish_Trabzon 0.256996 Basque_Spanish 0.256589 Chechen 0.256514 Icelandic 0.256418 Norwegian 0.256325 Lezgin 0.256272 Irish 0.256227 Tabasaran 0.256092 Italian_Bergamo 0.25605 English_Cornwall 0.256032 Polish_East 0.255991 Scottish 0.255955 Adygei 0.255913 Steppe_Maykop Latvian 0.261845 Russian_North 0.26145 Estonian 0.260355 Finnish 0.260211 Lithuanian 0.260072 Udmurd 0.259804 Ingrian 0.259663 Surui 0.259637 Vepsa 0.259608 Karelian 0.259532 Karitiana 0.259482 Russian_West 0.259397 Russian_Central 0.259274 Wichi 0.259106 Saami 0.258982 Komi 0.258945 Icelandic 0.258854 Swedish 0.258814 Mordovian 0.258604 Irish 0.25859Eyeballing the stats might be enough to get a general impression about what they mean, but to understand them properly it's necessary to get technical with something like PAST3 (see here). That's because f3-stats pick up shared genetic drift from all drift paths, and don't especially focus on more recently shared ancestry. This can often lead to confusing outcomes. Below are a few examples of linear models based on my f3-stats. Note that many Indo-European speakers, especially from Northern Europe, are foremost attracted to ancient samples from the Pontic-Caspian steppe. On the other hand, non-Indo-European speakers, from such far flung locations as the Caucasus and Iberia, show relatively stronger affinity to ancient samples from Anatolia and the Caucasus. Moreover, Uralic speakers show elevated affinity to ancient hunter-gatherer samples from Eastern Europe and Siberia. Makes sense, right? Based on these and other data, I'd say that Maykop and the culturally related Steppe Maykop were something of a multi-ethnic polity, with many near and far related languages spoken by its people, including perhaps Kartvelian, Northwest Caucasian, Yeniseian and Indo-European. But it seems to me that Proto-Indo-European was spoken by steppe foragers turned pastoralists just outside of the Maykop zone. And I'm quite sure that after the Maykop collapse various early Indo-European groups pushed across the Caucasus and deep into the Near East. Just take a look at the f3-stats and linear model for Hajji_Firuz_BA to see what I mean. See also... An exceptional burial indeed, but not that of an Indo-European The Steppe Maykop enigma Late PIE ground zero now obvious; location of PIE homeland still uncertain, but...
Labels:
Admixtools,
ancient DNA,
Bronze Age,
Caucasus,
f3-stats,
formal statistics,
Indo-European,
Maykop,
Meshoko,
migration,
PIE,
Piedmont Eneolithic,
Pontic-Caspian steppe,
Siberia,
Steppe Maykop,
Yamnaya
Wednesday, February 27, 2019
The Steppe Maykop enigma
Who were the Steppe Maykop people exactly? Their ancestry must surely rank as one of the biggest surprises served up by ancient DNA to date.
I always thought that they'd turn out roughly like a mixture between populations associated with the Kura-Araxes and Yamnaya cultures (mostly because their territory was located sort of in between them). Nope, that wasn't even close. This is where they cluster compared to Kura-Araxes and Yamnaya samples in my Principal Component Analysis (PCA) of world-wide genetic variation: the Global25.
To explore the ancestry of the Steppe Maykop people in more detail I ran a couple of unsupervised Global25/nMonte tests, using basically every ancient population in the (scaled) Global25 datasheet that seemed chronologically sensible and even remotely relevant. I narrowed things down to these two mixture models.
Steppe_Maykop Geoksiur_Eneolithic,11.2 Piedmont_Eneolithic,44.4 West_Siberia_N,44.4 distance%=1.5161 Steppe_Maykop Piedmont_Eneolithic,46.6 Sarazm_Eneolithic,10.4 West_Siberia_N,43 distance%=1.6408But, you might say, Global25/nMonte isn't a published analytical method and it doesn't run on formal statistics, the meat and potatoes of ancient DNA papers. OK then, let's try the same models with the qpAdm software, which is a published method and does run on formal statistics, using exactly the same samples.
Steppe_Maykop Geoksiur_Eneolithic 0.100±0.032 Piedmont_Eneolithic 0.433±0.053 West_Siberia_N 0.467±0.028 chisq 19.155 tail prob 0.159096 Full output Steppe_Maykop Piedmont_Eneolithic 0.429±0.051 Sarazm_Eneolithic 0.119±0.033 West_Siberia_N 0.452±0.026 chisq 18.090 tail prob 0.202699 Full outputThey're basically identical. Importantly, my models must reflect reality at some level, because otherwise I wouldn't be able to produce a pair of essentially identical results using such vastly different statistical methods. So the pertinent question is what do these results actually mean? It seems unlikely to me that we're dealing here with a highly complex three-way mixture process, involving populations from such far flung locations as western Siberia and southern Central Asia. Rather, I suspect that Steppe Maykop was the result of a two-way mixture between Piedmont_Eneolithic (the population that lived before it on the steppe north of the Caucasus) and someone just a little bit more easterly. I'm guessing that the latter was the (as yet unsampled) population associated with the Kelteminar archeological culture. By the way, please note that Piedmont_Eneolithic is made up of samples from two different locations on the Piedmont steppe, and I occasionally treat them as separate populations labeled Progress_Eneolithic and Vonyuchka_Eneolithic (for instance, see here). Update 28/02/2019: Below is a PCA focusing on West Eurasian genetic variation. Overall, the position of Steppe Maykop relative to Geoksiur_Eneolithic, Piedmont_Eneolithic and West_Siberia_N appears to reflect my nMonte and qpAdm models. However, as per our discussion in the comments, one of the Steppe Maykop individuals (the most southerly one in the PCA) probably also has recent ancestry from the Caucasus. See also... An exceptional burial indeed, but not that of an Indo-European Maykop: a multi-ethnic layer cake? Late PIE ground zero now obvious; location of PIE homeland still uncertain, but...
Labels:
admixture,
ancient DNA,
Bronze Age,
Caucasus,
Global25,
Maykop,
migration,
Piedmont Eneolithic,
Pontic-Caspian steppe,
Progress Eneolithic,
qpAdm,
Steppe Maykop,
Vonyuchka Eneolithic,
West Siberia,
Yamnaya
Saturday, February 23, 2019
Catacomb > Armenia_MLBA
It's now clear, thanks to ancient DNA, that Transcaucasia and surrounds were affected by multiple, and at times significant, population movements from Eastern Europe during the Chalcolithic and Bronze Age periods. Based on the ancient samples from what is now Armenia, I'd say that this process peaked during the Middle Bronze Age. But who exactly were the people who perhaps swarmed south of the Caucasus at this time?
The most likely suspects are the various groups that occupied the southernmost parts of the Pontic-Caspian steppe throughout the Bronze Age. They were associated with the so called Catacomb, Kubano-Tersk and Yamnaya archeological cultures. Below is a Principal Component Analysis (PCA) that compares samples from these cultures with those from Middle to Late Bronze Age Armenia (labeled Armenia_MLBA). The relevant datasheet is available here.
Note that Armenia_MLBA forms a cline that appears to be stretching out towards the Catacomb, Kubano-Tersk, Yamnaya and other Bronze Age steppe groups, and this suggests that it harbors significant and probably recent steppe-related ancestry. But PCA plots based on just two dimensions of genetic variation can be misleading at times, so let's check this out with some formal mixture models using qpAdm.
Armenia_MLBA Catacomb 0.234±0.028 Kura-Araxes_Kaps 0.766±0.028 chisq 10.723 tail prob 0.826248 Full output Armenia_MLBA Kubano-Tersk 0.254±0.030 Kura-Araxes_Kaps 0.746±0.030 chisq 13.535 tail prob 0.633284 Full output Armenia_MLBA Kura-Araxes_Kaps 0.768±0.028 Yamnaya_Kalmykia 0.232±0.028 chisq 14.454 tail prob 0.564954 Full output Armenia_MLBA Kura-Araxes_Kaps 0.762±0.029 Yamnaya_Caucasus 0.238±0.029 chisq 15.916 tail prob 0.458816 Full outputAll of these models are statistically very sound, and even though I ranked the results by "tail prob", there's nothing in the output that clearly points to any one of the southern steppe groups as the obvious source of the steppe-related ancestry in Armenia_MLBA. But, interestingly, Catacomb tops the ranking, and it probably also makes the most sense based simply on Carbon-14 chronology. So, for now, I'm going with Catacomb. I didn't get a chance yet to investigate this issue in detail with the Global25. Does it contradict the results from my PCA and qpAdm analyses? If anyone reading this would like to take a close look that'd be great. Feel free to post your findings in the comments below. And if the answer is indeed Catacomb, then what language did these Catacomb-derived migrants, or perhaps invaders, speak? If not proto-Armenian then what? By the way, please be aware that the Kubano-Tersk samples in my analyses are the same individuals as those featured in Wang et al. 2019 under the label "North Caucasus". See also... Early chariot drivers of Transcaucasia came from... Likely Yamnaya incursion(s) into Northwestern Iran Late PIE ground zero now obvious; location of PIE homeland still uncertain, but...
Labels:
Anatolia,
ancient DNA,
Armenia,
Bronze Age,
Catacomb,
Caucasus,
Eastern Europe,
Greco-Armenian,
Hittites,
Indo-European,
Kubano-Tersk,
Pontic-Caspian steppe,
Transcaucasia,
Yamna,
Yamnaya
Friday, April 27, 2018
The mystery of the Sintashta people
During the Middle to Late Bronze Age, the steppes southeast of the Ural Mountains, in what is now Russia, were home to communities of metallurgists who buried their warriors with horses and the earliest examples of the spoked-wheel battle chariot.
We don't know what they called themselves, because they didn't leave any written texts, but their archaeological culture is commonly known as Sintashta. It was named after a river near one of their main settlements; an elaborate fortified town that has also been described as an ancient metallurgical industrial center. Another of their well known settlements, very similar to Sintashta, is Arkaim, pictured below courtesy of Wikipedia.
Sintashta is arguably one of the coolest ancient cultures ever discovered by archaeologists. It's also generally accepted to be the Proto-Indo-Iranian culture, and thus linguistically ancestral to a myriad of present-day peoples of Asia, including Indo-Aryans and Persians. No wonder then, that its origin, and that of its population, have been hotly debated issues.
The leading hypothesis based on archaeological data is that Sintashta is largely derived from the more westerly and warlike Abashevo culture, which occupied much of the forest steppe north of the Black and Caspian Seas. In turn, Abashevo is usually described as an eastern offshoot of the Late Neolithic Corded Ware Culture (CWC), which is generally seen as the first Indo-European archaeological culture in Northern Europe (see here).
Below is a Principal Component Analysis (PCA) featuring 38 Sintashta individuals from the recent Narasimhan et al. 2018 preprint. Note that the main Sintashta cluster overlaps almost perfectly with the main CWC cluster. The relevant datasheet is available here.
Moreover, many ancient and present-day South and Central Asians, particularly those identified with or speaking Indo-Iranian languages, appear to be strongly attracted to the main Sintashta cluster, forming an almost perfect cline between this cluster and the likely Indus Valley diaspora individuals who show no evidence of steppe ancestry.
This is in line with mixture models based on formal statistics showing significant Sintashta-related ancestry in Indo-Iranian-speakers (for instance, see here), and high frequencies of Y-haplogroup R1a-Z93 in both the Sintashta and many Indo-Iranian-speaking populations.
Some of the Sintashta samples are outliers from the main Sintashta cluster, and that's because they harbor elevated levels of ancestry related to the Mesolithic and Neolithic foragers of Eastern Europe and/or Western Siberia. This is especially true of a pair of individuals who belong to Y-haplogroup Q. However, this doesn't contradict archaeological data, which suggest that the Sintashta community may have been multi-cultural and multi-lingual. Indeed, it's generally accepted based on historical linguistics data that there were fairly intense contacts in North Eurasia between the speakers of Proto-Indo-Iranian, Proto-Uralic and Yeniseian languages.
Thus, it appears that there's not much left to debate because ancient DNA has seemingly backed up the most widely accepted hypotheses about the origin of Sintashta and its people, and their identification mainly as Proto-Indo-Iranian-speakers.
However, a sample from a Sredny Stog II culture burial on the North Pontic steppe, in what is now eastern Ukraine, has complicated matters somewhat. This individual, known as Ukraine_Eneolithic I6561, not only clusters very strongly with the most typical Sintashta samples, but also belongs to Y-haplogroup R1a-Z93. On the other hand, none of the CWC remains sequenced to date belong to this particular subclade of R1a (although, obviously, they do belong to a host of near and far related R1a subclades).
I've never seen anyone worth reading propose that Sintashta might derive from Sredny Stog II instead of Abashevo. And no wonder, because Sredny Stog II was long gone when Sintashta appeared in the archaeological record.
But if CWC remains continue to fail to produce R1a-Z93, while, at the same time, the steppes of eastern Ukraine and surrounds are shown to be a hotbed of R1a-Z93 from the Sredny Stog to the Sintashta periods, which I think is possible, then ancient DNA might well force a serious re-examination of how the awesome Sintashta culture and people came to be.
See also...
Subscribe to:
Posts (Atom)




















