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Showing posts with label Yamna. Show all posts
Showing posts with label Yamna. Show all posts

Sunday, June 22, 2025

‘Proto-Yamnaya’ Eneolithic individuals from Kuban steppe c. 3700 BC ? (guest post)


This is a guest post by an anonymous contributor. I don't necessarily agree with its findings, but I think it's a good way to get the ball rolling here again. Feel free to let me know what you think. Please note, however, that any comments that show mental instability will be blocked. No more crazy talk on this blog.

In order to understand who Yamnaya people were, one must first define ‘Yamnaya’. We will adopt a strictu sensu view (e.g. Anthony, Heyd) encompassing burials dating 3200-2600 BC, with a characteristic body position, mound construction, and copper artefacts. These complexes can be linked to a core group of people whose autosomal make-up is quite homogeneous throughout their wide geographic range. Moreover, almost all males belong to Y-haplogroup R1b-M269-Z2103. In this light, ‘core Yamnaya’ does not represent a ‘proto-Indo-European’ population, as commonly proclaimed, but a group which contributed to several post-PIE population-language complexes, such as Tocharian, Armenian and some Paleo-Balkan languages. However, historical linguistics is not the focus of this post.

Archeologists had linked Yamnaya to earlier complexes such as Khvalynsk, Repin and/or Mikhalivka. Given that cultural markers such as pottery and burial customs can be borrowed and copied, ancient DNA can offer a more objective assessment of population origins. However, the cacophony of clusters, clines and other statistical constructs in publications can be confusing. A more rationalized approach is required, and one way is to co-analyse phylogenetically linked individuals across space and time. Apart from a lower-quality individual from Smyadovo (Bulgaria c. 4300 BC), the earliest attestation of R1b-M269 is in two individuals from the Kuban steppe (Stavropol region) c. 3700 BC -NV3003 and KST001 (Ghaliachi et al 2024). However, Y-hg R1b-M269 is missing in currently sampled Kuban steppe and north Caucasian males from the preceding period (5000-4000 BC). Males of the ‘Kuban steppe 4500bc’ group (Progress, Vojnucka, Sengeleevskiy, etc) are instead derived for the phylogenetically divergent Y-hg Rb-V1636. Males from the Nalchik cemetery are also derived for Y-hg R1b-V1636, or related haplotypes, although they were buried in a ‘Caucasian Farmer’ pose and heavily infused with such ancestry, but probably also had a burial mound thrown above. We do not know when the R1b-V1636 clans entered the northern Caucasus region, or from where, but they appear to have been attracted by trade with North Caucasian Famer (~Eneolithic) groups- termed as ‘Meshoko-Zamok’, ‘Chokh’, etc, in literature. Curiously the Nalchik group has minimal Central Asian (“TTK-related”) ancestry, whilst the Kuban steppe group has high levels. This suggests that TTK-related ancestry arrived after R1b-V1636 dominant EHG clans entered the North Caucasus region, but other scenarios are possible. Lastly, two ‘Meshoko culture’ males from Unakozovskaya have been assigned to Y-hg J2a-L26.

A shake-up occurred in the north Caucasus after 4000 BC. As we know, this corresponds to the emergence of the Majkop phenomenon, catalysed by renewed migrations from the south. These were not ‘Uruk migrants’ as sometimes proposed - the Uruk phenomenon occurred several hundred years later and was a south Mesopotamian phenomenon. Instead, these newcomers emerge from southern Caucasus- north Mesopotamian ‘Late Chalcolithic’ groups. They brought with them multiple West Asian lineages, such as Y-hg T, L2, J2a-, J2b, G2. Over time they mixed with preceding north Caucasian Eneolithic groups, culminating in the Novosvobodnaja phenomenon.

The emergence of Majkop as a new socio-cultural complex broke down the previous system dominated by Y-hg R1b-V1636 clans. The Majkop sphere consisted of a ‘core’ of heterarchical chiefs buried in elaborate kurgans near the Mountains, and a dynamic northern ‘frontier’ in the steppe lands (as far as the lower Don) between 4000 and 3000 BC. At least 3 ‘‘Majkop periphery’ genetic groups can be defined; in fact all these groups can be termed ‘steppe Majkop’:

1- Group with western Siberian/ north central Asian ancestry (the ‘genetic steppe Majkop’ as defined in Wang et al, 2023)
2- The South Caucasian/north Iranian ‘Zolotarevka’ group
3- The R1b-M269 duo.

Regardless of their lineages and genomic affinities, these individuals were often buried in kurgans which over time formed groups. These were not continuations of pre-4000 BC kurgans, but the communities instead made a conscious choice to build new kurgans after 4000 BC, adding to the idea of discontinuity. But once built, these kurgan clusters continued to be developed for hundreds of years, into the Yamnaya period. This does imply ethnic homogeneity or continuity, just a ‘continuity of place’. Without a direct attestation of a phylogenetic ancestor, and guestimating from their (non-identical) genomic profile, we are left to speculate that Y-hg R1b-M269 individuals moved down from somewhere in the Volga-Don interfluve. Perhaps amongst groups utilizing Repin pottery, but if so, they did not continue its use in their new contexts. By 3000 BC, the Majkop system collapsed. Yamnaya groups and their ‘Catacomb’ descendants took control of the north Caucasus region, having benefitted from years of trade/ exchange and knowledge gathering. Whether Yamnaya actually descend from individuals like KST-1 or NV3003 remains to be seen, however these are the closest leads we have. Certainly, we can model Yamnaya as deriving from KST-1 (88%) + Dnieper_N (12%), but we should be cautious when using singular individuals as ‘sources’.
See also...

The PIE homeland controversy: December 2024 open thread

Thursday, September 1, 2022

Dear Iosif #3


Back in 2016 I made this prediction about the origins of the Yamnaya people (Steppe_EMBA):

But here's my prediction: Steppe_EMBA only has 10-15% admixture from the post-Mesolithic Near East not including the North Caucasus, and basically all of this comes via female mediated gene flow from farming communities in the Caucasus and perhaps present-day Ukraine.

The relevant blog post is still here. Looking back, my analysis is a bit sloppy and I didn't articulate my ideas too well. But that was a pretty good prediction for its time, and I believe it still has a chance of being confirmed, more or less.

On the other hand, the widely publicized hypothesis that the Yamnaya population is a ~50/50 mixture between indigenous Eastern European hunter-gatherers and Near Eastern or West Asian migrants never looked right to me. So I'm glad that it's now dead and buried.

Those of you not up to date with this topic, all you need to know is that the Yamnaya genotype existed in Eastern Europe at least a thousand years before Yamnaya, and, moreover, the Yamnaya people are largely derived from Eastern European foragers already rich in Near Eastern-related ancestry. The relevant ancient genomes are on the way (for instance, see here).

Nevertheless, the narrative that waves of Near Eastern migrants moved into prehistoric Eastern Europe, leading to the emergence of the Yamnaya culture and even the Proto-Indo-European language, is still being pushed by some notable scientists working with ancient DNA.

My hope is that, considering the latest revelations about the genetic origins of the Yamnaya people, these scientists can embrace a more nuanced view. How about something like this?

- people moved around, and they were especially mobile on the Eastern European steppe from the Eneolithic onwards

- when they made contact they sometimes mixed, so there was admixture between far flung steppe groups

- since population densities on the steppe were low until the Yamnaya period, minor admixture that entered the steppe during the Neolithic and Eneolithic wasn't dilluted easily.

See also...

Dear Iosif #2

But Iosif, what about the Phrygians?

Dear Iosif, about that ~2%

Dear Iosif...Yamnaya

Thursday, February 10, 2022

Mainstream media BS: Europeans owe their height to Asian nomads


From a recent Daily Mail article by some clown named Sam Tonkin:

Present day Europeans owe their blue eyes to hunter gatherers, their height to Asian nomads and their blonde hair to Anatolian Neolithic farmers, a new study suggests.

...

Most of the contemporary European genetic makeup was shaped by movements that occurred in the last 10,000 years when local hunter gatherers mixed with incoming Anatolian farmers — from present-day Turkey — and Asian nomads, or Pontic Steppe pastoralists.

The latter originated from what is now parts of Bulgaria, Romania, Moldova, Ukraine, Russia and Kazakhstan.

Haha.

Bulgaria, Romania, Moldova and Ukraine are European countries. The relevant parts of Russia and Kazakhstan are also located in Europe.

Obviously, the author is referring to the Yamnaya herders who lived on the Pontic-Caspian steppe, which is obviously in Eastern Europe.

I blame Johannes Krause for this.

See also...

Matters of (basic) geography

Blond hair is only indirectly associated with Anatolian ancestry in Estonia...duh

Ancient ancestry and complex traits in Estonians (Marnetto et al. 2022)

Sunday, January 23, 2022

Para-Turbo-Balto-Slavic?


I'm seeing increasing numbers of Bronze and Iron Age samples from Central Europe and surrounds with this peculiar set of traits:

- shared genetic drift with present-day Balto-Slavic speakers to the exclusion of most other Europeans

- and yet, an unusually low level of Yamnaya-related steppe ancestry

- so much so, in fact, that they're often outside the range of modern European genetic variation.

As far as I can tell, currently the best examples of this unusual population are HUN_Mako_EBA_o:I1502 (Mathieson et al. Nature 2015) and HUN_EIA_Prescythian_Mezocsat_o1:I18241 (Patterson et al. Nature 2021). Both are from the Carpathian Basin in what is now Hungary.

I ran a series of qpAdm mixture models to try and learn more about their origins. The most robust outcomes, out of about 50 different attempts, are these:

right pops:
CMR_Shum_Laka_8000BP
MAR_Taforalt
IRN_Ganj_Dareh_N
Levant_PPNB
TUR_Barcin_N
Iberia_Southeast_Meso
UKR_Meso
England_Meso
RUS_Karelia_HG
RUS_West_Siberia_HG
MNG_North_N
TWN_Hanben
BRA_LapaDoSanto_9600BP

HUN_Mako_EBA_o
Baltic_LTU_Narva 0.149 ∓0.028
POL_Globular_Amphora 0.613 ∓0.028
Yamnaya_RUS_Samara 0.238 ∓0.029
chisq 10.836
tail prob 0.370463
Full output

HUN_EIA_Prescythian_Mezocsat_o1
Baltic_LTU_Narva 0.186 ∓0.028
POL_Globular_Amphora 0.592 ∓0.027
Yamnaya_RUS_Samara 0.222 ∓0.029
chisq 12.492
tail prob 0.253499
Full output

Combining the two genomes produces a very similar result:

HUN_EBA-EIA_o
Baltic_LTU_Narva 0.160 ∓0.023
POL_Globular_Amphora 0.612 ∓0.023
Yamnaya_RUS_Samara 0.227 ∓0.023
chisq 14.653
tail prob 0.14524
Full output

Importantly, when I move RUS_Karelia_HG from the right pops to the left pops, to test whether HUN_EBA-EIA_o really has steppe ancestry, as opposed to closely related hunter-gatherer ancestry, I still get a very similar outcome:

HUN_EBA-EIA_o
Baltic_LTU_Narva 0.158 ∓0.027
POL_Globular_Amphora 0.605 ∓0.033
RUS_Karelia_HG 0.014 ∓0.038
Yamnaya_RUS_Samara 0.223 ∓0.053
chisq 10.461
tail prob 0.234171
Full output

So these largely Globular Amphora-related individuals do harbor as much as a quarter of steppe ancestry, which is to be expected considering the massive genetic turn-over that most of Europe experienced just before their time as a result of population expansions from the Pontic-Caspian steppe.

Nevertheless, this is ~20% less steppe ancestry than in the present-day populations of the region, and it clearly shows in any decent Principal Component Analysis (PCA) of West Eurasia. For instance:
At the same time, the relatively close genetic relationship between these ancients and present-day Balto-Slavic speaking populations shows up in fine-scale intra-European PCA.

The origins and implications of this population are still a mystery to me. I don't think it's native to the Carpathian Basin. Indeed, my qpAdm models suggest that it may have moved into this region from somewhere to the northeast, because its ancestry is best modeled with ancient groups from present-day Lithuania, Poland and Russia.

I'm adamant that these people weren't Balto-Slavic speakers, and certainly not proto-Slavs. Rather, I suspect that much like the Welzin warriors of Bronze Age North-Central Europe, they were closely related to a contemporaneous group that eventually gave rise to proto-Slavs. At best, they may have somehow contributed to the ethnogenesis of Balto-Slavs.

By the way, using the Global25 to model their ancestry is highly problematic, because of the strong Balto-Slavic genetic drift that affects some of the dimensions. So be careful when you try it, or better yet, don't try it at all, and stick to formal stats in this particular instance.

See also...

Tollense Valley Bronze Age warriors were very close relatives of modern-day Slavs

Friday, January 21, 2022

Yamnaya is from Europe, but it's really from Asia


I was about to post a comment under a new preprint at bioRxiv, but the comment section isn't there anymore. Hopefully, this is just a temporary glitch.

The preprint in question is titled Reconstructing the spatiotemporal patterns of admixture during the European Holocene using a novel genomic dating method [LINK]. It's co-authored by Harvard/Broad MIT scientist Nick Patterson who occasionally comments at this blog.

My impression is that the authors see the people associated with the Yamnaya culture as Asians who simply used "far" Eastern Europe as a springboard to expand into other parts of Europe.

If so, they're dead wrong.

There are at least three arguments why the Yamnaya population should be seen as quintessentially European:

- its home was initially and overwhelmingly the Pontic-Caspian steppe, which is entirely located within the present-day borders of Europe

- Yamnaya genomes are clearly different from those of older populations native to nearby parts of Asia, and, in fact, these differences show a very strong correlation with the present-day borders between Europe and Asia

- the Yamnaya people weren't a new population in Europe by any stretch, but must have been overwhelmingly derived from the very similar Eneolithic peoples of the Pontic-Caspian steppe and/or the nearby forest steppe, both of which are located in Eastern Europe.

And yet, this is what the preprint claims:

The beginning of the Bronze Age was a period of major cultural and demographic change in Eurasia, accompanied by the spread of Yamnaya Steppe Pastoralist-related ancestry from Pontic-Caspian steppes into Europe and South Asia (16).

In fact, what really happened at this time was that Yamnaya steppe pastoralist-related ancestry spread from Eastern Europe to other parts of Europe, as well as to Central and West Asia.

The preprint does eventually explain that present-day South Asians derive their Yamnaya-related ancestry from a later eastward expansion of the European Corded Ware culture (CWC), but it completely ignores the fact that the Afanasievo culture was the result of the initial eastward expansion from Europe to Asia. That is, the ancestors of the Afanasievo people were recent migrants from the Pontic-Caspian steppe to Central Asia and Siberia.

There's also this:

Over the following millennium, the Yamnaya-derived groups of the Corded Ware Complex (CWC) and Bell Beaker complex (BBC) cultures brought Steppe pastoralist-related ancestry to Europe.

Seriously? Both the CWC and BBC, just like the Yamnaya culture, were from Europe. In fact, as per above, the descendants of the CWC expanded into Asia.

And this:

The second major migration occurred when populations associated with the Yamnaya culture in the Pontic-Caspian steppe expanded to central and western Europe from far eastern Europe.

The authors basically admit here that Yamnaya came from Eastern Europe, but they call it "far" Eastern Europe. Perhaps they know something I don't, but as things stand, there's no evidence that Yamnaya came from "far" Eastern Europe. In fact, the emerging consensus based on ancient DNA, including pre-publication data, is that Yamnaya may have originated in what is now Ukraine. In my opinion, Ukraine isn't located in "far" Eastern Europe, but more or less in the middle of it.

Inexplicably, this is what they say about the genetic origins of the Yamnaya and Afanasievo peoples:

These groups were likely the result of a genetic admixture between the descendants of EHG-related groups and CHG-related groups associated with the first farmers from Iran (8, 22, 36).

...

Thus, we combined all early Steppe pastoralist individuals in one group to obtain a more precise estimate for the genetic formation of proto-Yamnaya of ~4,400 to 4,000 BCE (Figure 2). These dates are noteworthy as they pre-date the archeological evidence by more than a millennium (37) and have important implications for understanding the origin of proto-Pontic Caspian cultures and their spread to Europe and South Asia.

Not really.

Like I said, the Yamnaya population was overwhelmingly derived from the Eneolithic peoples of the Eastern European steppe and/or forest steppe. And these Yamnaya-like Eneolithic peoples were spread out across a vast area of Eastern Europe by at least ~4,500 BCE. Some of their genomes have been available for several years, and many more are on the way.

It is possible that the Yamnaya and Afanasievo genotype formed in 4,400-4,000 BCE, but if so, then this was due to mixing between the Eneolithic steppe peoples and nearby European farmers. That's because the difference between the Yamnaya and Eneolithic steppe genotypes is minor (~15%) European farmer admixture in the former.

The really interesting puzzle is exactly where and when the peculiar Eneolithic steppe genotype came into being. Any ideas Dr Patterson?

See also...

Matters of geography

Understanding the Eneolithic steppe

Wednesday, September 15, 2021

Yamnaya people drank horse milk (Wilkin et al. 2021)


Over at Nature at this LINK. I'm guessing the claim that Yamnaya pastoralists lived in Scandinavia is a huge typo. Obviously, the authors are referring to the people of the Corded Ware culture (CWC). From the paper:

During the Early Bronze Age, populations of the western Eurasian steppe expanded across an immense area of northern Eurasia. Combined archaeological and genetic evidence supports widespread Early Bronze Age population movements out of the Pontic–Caspian steppe that resulted in gene flow across vast distances, linking populations of Yamnaya pastoralists in Scandinavia with pastoral populations (known as the Afanasievo) far to the east in the Altai Mountains1,2 and Mongolia3. Although some models hold that this expansion was the outcome of a newly mobile pastoral economy characterized by horse traction, bulk wagon transport4,5,6 and regular dietary dependence on meat and milk5, hard evidence for these economic features has not been found. Here we draw on proteomic analysis of dental calculus from individuals from the western Eurasian steppe to demonstrate a major transition in dairying at the start of the Bronze Age. The rapid onset of ubiquitous dairying at a point in time when steppe populations are known to have begun dispersing offers critical insight into a key catalyst of steppe mobility. The identification of horse milk proteins also indicates horse domestication by the Early Bronze Age, which provides support for its role in steppe dispersals. Our results point to a potential epicentre for horse domestication in the Pontic–Caspian steppe by the third millennium bc, and offer strong support for the notion that the novel exploitation of secondary animal products was a key driver of the expansions of Eurasian steppe pastoralists by the Early Bronze Age.

Wilkin, S., Ventresca Miller, A., Fernandes, R. et al. Dairying enabled Early Bronze Age Yamnaya steppe expansions. Nature (2021). https://doi.org/10.1038/s41586-021-03798-4

See also...

On the origin of the Corded Ware people

Tuesday, July 20, 2021

On the origin of the Corded Ware people


There's been a lot of talk lately about the finding that the peoples associated with the Corded Ware and Yamnaya archeological cultures were genetic cousins (for instance, see here). As I've already pointed out, this is an interesting discovery, but, at this stage, it's difficult to know what it means exactly.

It might mean that the Yamnayans were the direct predecessors of the Corded Ware people. Or it might just mean that, at some point, the Corded Ware and Yamnaya populations swapped women regularly (that is, they practiced female exogamy with each other).

In any case, I feel that several important facts aren't being taken into account by most of the interested parties. These facts include, in no particular order:

- despite being closely related, the Corded Ware and Yamnaya peoples were highly adapted to very different ecological zones - temperate forests and arid steppes, respectively - and this is surely not something that happened within a few years and probably not even within a couple of generations

- both the Corded Ware and Yamnaya populations expanded widely and rapidly at around the same time, but never got in each others way, probably because they occupied very different ecological niches

- despite sharing the R1b Y-chromosome haplogroup, their paternal origins were quite different, with Corded Ware males rich in R1a-M417 and R1b-L51 and Yamnaya males rich in R1b-Z2103 and I2a-L699

I suppose it's possible that the Corded Ware people were overwhelmingly and directly derived from the Yamnaya population. But right now my view is that, even if they were, then the Yamnaya population that they came from was quite different from the classic, R1b-Z2103-rich Yamnaya that spread rapidly across the steppes.

Indeed, perhaps what we're dealing with here is a very early (proto?) Yamnaya gene pool located somewhere in the border zone between the forests and the steppes, that then split into two main sub-populations, with one of these groups heading north and the other south?

I do wonder what David Anthony would say if he was made aware of the above mentioned facts? Then again, perhaps he's already aware of them, and simply chose to ignore them when formulating his latest theory about the origin of the Corded Ware people?

See also...


Monday, June 28, 2021

The PIE homeland controversy: June 2021 status report


Archeologist David Anthony has made several appearances online recently to promote his theories about the origins of the Corded Ware and Yamnaya cultures and peoples.

In a clip on Youtube he reiterated his theory that the so called Iranian-related ancestry in the Yamnaya people actually came from what is now Iran, and, more precisely, that it was carried by hunter-gatherers who travelled relatively rapidly from the South Caspian region into the Volga Delta in what is now Russia.

It's still a complete mystery to me as to why a group of hunter-gatherers from the South Caspian would undertake such a migration, instead of, say, expanding their range gradually over thousands of years, first into the Caucasus and eventually into Eastern Europe.

But there's a more serious problem with Anthony's theory: it contradicts the currently available ancient DNA. That's because the so called Iranian-related ancestry in the Yamnaya people is most closely related to the Kotias and Satsurblia hunter-gatherers from what is now Georgia, and these hunter-gatherers form a separate clade from the earliest samples from what is now Iran. For instance, see here and here.

Also, in a podcast on Razib's blog, Anthony doubled down on his theory that Y-chromosome haplogroup R1a was closely associated with Yamnaya plebs who were excluded from Kurgan burials, and, as a result, their remains haven't yet been sampled.

At least this theory isn't yet contradicted by ancient DNA, but it's more complicated and less parsimonious than my theory, which posits that R1a, or rather R1a-M417, was simply a very rare lineage in the Yamnaya population, and that it only became a common and widespread marker thanks to the Corded Ware expansion (see here).

Intriguingly, my understanding is that there are several unpublished R1a samples from the Caspian and Volga steppes at Harvard's David Reich Lab that have been classified by its scientists as Yamnaya outliers. Of course, Anthony is collaborating on at least one major paper with this lab (see here).

Ergo, I strongly suspect that Anthony's theory is in part based on these Yamnaya outliers. However, I also believe that these samples are wrongly dated and probably represent Scythians and/or Sarmatians. I'll be able to look into that if they're ever published.

Speaking of the David Reich Lab, its leading scientists, David Reich and Nick Patterson, have also made appearances online recently, on Youtube and Razib's blog, respectively, to reveal that the Corded Ware and Yamnaya peoples aren't just very similar genetically, but in fact close cousins.

This is a very interesting finding. Apparently it's based on a relatively high level of Identity-by-Descent (IBD) segment sharing between Corded Ware and Yamnaya samples, but that's all I know. I'm guessing that the relevant paper is coming soon (that is, within the next five years).

However, the long-standing question that the readers of this blog want to see answered is not whether the Corded Ware and Yamnaya peoples are close cousins, but whether Yamnaya migrants founded the Corded Ware culture. The obvious way to prove that they did is to find at least one ancient population unambiguously classified as part of the Yamnaya horizon that is rich in the typically Corded Ware Y-haplogroups R1a-M417 and R1b-L151.

See also...

On the origin of the Corded Ware people

The PIE homeland controversy: January 2019 status report

The PIE homeland controversy: August 2019 status report

Thursday, February 13, 2020

Ancient DNA vs Ex Oriente Lux


In recent years you may have read academic papers, books and press articles claiming that the Early Bronze Age Yamnaya culture of the Pontic-Caspian steppe was founded by migrants from the Caucasus, Mesopotamia or even Central Asia.

Of course, none of this is true.

The Yamnaya herders and closely related groups, such as the people associated with the Corded Ware culture, expanded from the steppe between the Black and Caspian seas, and, thanks to ancient DNA, it's now certain that they were overwhelmingly derived from a population that had existed in this region since at least the mid-5th millennium BCE (see here).

So rather than being culturally advanced colonists from some Near Eastern civilization, the ancestors of the Yamnaya herders were a relatively primitive local people who still largely relied on hunting and fishing for their subsistence. They also sometimes buried their dead with flint blades and adzes, but hardly ever with metal objects, despite living in the Eneolithic epoch or the Copper Age.

As far as I know, this group doesn't have a specific name. But in recent scientific literature it's referred to as Eneolithic steppe, so let's use that.

It's not yet clear how the Yamnaya people became pastoralists. Some scholars believe that they were basically an offshoot of the cattle herding Maykop culture of the North Caucasus. However, the obvious problem with this idea is that the Yamnaya and Maykop populations probably didn't share any recent ancestry. In fact, ancient DNA shows that the former wasn't derived from the latter in any important or even discernible way (see here).

On the other hand, Yamnaya samples do harbor a subtle signal of recent gene flow from the west that appears to be most closely associated with Middle to Late Neolithic European agropastoralists (see here). Therefore, it's possible that herding was adopted by the ancestors of the Yamnaya people as a result of their sporadic contacts with populations living on the western edge of the Pontic-Caspian steppe.

Eneolithic steppe is currently represented by just three samples in the ancient DNA record, and all of these individuals are from sites on the North Caucasus Piedmont steppe (two from Progress 2 and one from Vonyuchka 1).

As a result, it might be tempting to argue that cultural, if not genetic, impulses from the Caucasus did play an important role in the formation of the Yamnaya and related peoples. However, it's important to note that the North Caucasus Piedmont steppe was the southern periphery of Eneolithic steppe territory.

Below is a map of Eneolithic steppe burial sites featured in recent scientific literature. It's based on data from Gresky et al. 2016, a paper that focused on a specific and complex type of cranial surgery or trepanation often practiced by groups associated with this archeological culture (see here).


Incredibly, one of the skeletons from Vertoletnoe pole has been radiocarbon dated to the mid-6th millennium BCE. My suspicion, however, is that this result was blown out by the so called reservoir effect (see here). In any case, the academic consensus seems to be that the roots of Eneolithic steppe should be sought in the Lower Don region, rather than in the Caucasus foothills (see page 36 here).

Considering that nine Eneolithic steppe skulls from the Lower Don were analyzed by Gresky et al., I'd say it's only a matter of time before we see the publication of genome-wide data for at least of couple of these samples. Indeed, the paper's lead author is from the Deutsches Archäologisches Institut, which is currently involved in a major archaeogenetic project on the ancient Caucasus and surrounds. Unfortunately, the study is scheduled to be completed in about four years (see here).

But whatever happens, the story of Eneolithic steppe deserves to be investigated in as much detail as possible, because it obviously had a profound impact on Europe and its people.

In my estimation, at least a third of the ancestry of present-day Northern Europeans, all the way from Ireland to the Ural Mountains in Russia, is ultimately derived from Eneolithic steppe groups. It's also possible that R1a-M417 and R1b-L51, the two most frequent Y-chromosome haplogroups in European males today, derive from a couple of Eneolithic steppe founders. If so, that's a very impressive effort for such an obscure archeological culture from what is generally regarded as a peripheral part of Europe.

See also...


Monday, December 30, 2019

A final note for the year


I feel like I've spent a good part of 2019 banging my head against a thicker than average brick wall.

Much of this feeling is tied to the controversy over the ethnogenesis of the Yamnaya people, and my often futile attempts to explain that their origin cannot be sought in what is now Iran, or, indeed, anywhere outside of Eastern Europe.

This post is my final attempt to lay out the facts in regards to this topic. Next year I'll have better things to do than to argue the bleeding obvious.

Below are two graphs from a Principal Component Analysis (PCA) based on relatively high quality ancient human genotype data from the Caucasus and surrounds. They include two typical Yamnaya individuals from burial sites north of the Caspian Sea. I made the graphs with the Vahaduo Custom PCA tool here. The relevant datasheet can be downloaded here.



Here's what I'm seeing:

- the Yamnaya individuals sit on genetic clines made up of hunter-gatherers native to the Caucasus and various parts of Eastern Europe, including a trio from the southernmost part of the Pontic-Caspian steppe (labeled Steppe_Eneolithic), with whom they form a distinct cluster

- the samples from the Caucasus and the Iranian Plateau form very different clusters, so there's no support here for the ancient Caucasus/Iranian grouping that is often haphazardly invoked in scientific literature

- there's no indication that the Yamnaya and/or Steppe_Eneolithic groups experienced recent gene flow, or, for that matter, any gene flow whatsoever, from what is now Iran.

Of course, analyses based on formal statistics suggest that the Yamnaya population harbors minor western ancestry that is missing in Steppe_Eneolithic. In fact, I was first to argue this point (see here). So let's add a couple of ancient farmers from Western Europe to my PCA to see how they affect the graphs. The relevant datasheet is available here.



Yep, the Yamnaya pair appears to be peeling away very slightly, but deliberately, from the Steppe_Eneolithic individuals towards the part of the plot occupied by the farmers.

Admittedly, I'm no Sherlock Holmes, but even with my fairly average sleuthing abilities, I'm pretty sure I know how the Yamnaya people came to be. They formed largely on the base of a population very much like Steppe_Eneolithic somewhere deep in Eastern Europe, well to the north of the Caucasus, and nowhere near the Iranian Plateau.

See also...

A note on Steppe Maykop

Thursday, September 26, 2019

Is Yamnaya overrated?


Four years after the publication of the seminal ancient DNA paper Massive migration from the steppe is a source for Indo-European languages in Europe by Haak et al., we're still waiting for some of its loose ends to be finally tied up with new samples. In particular...

- if the men of the Corded Ware culture (CWC) were, by and large, derived from the population of the Yamnaya culture, then where are the Yamnaya samples with R1a-M417, the main CWC Y-haplogroup?

- if the men of the Bell Beaker culture (BBC) were also, by and large, derived from the population of the Yamnaya culture, then where are the Yamnaya samples with R1b-P312, the main BBC Y-haplogroup?

- and, most crucially, if R1b-L51, which includes R1b-P312, and is nowadays by far the most important Y-haplogroup in Western Europe, arrived there from the Pontic-Caspian steppe, then why hasn't it yet appeared in any of the ancient DNA from this part of Eastern Europe or surrounds, except of course in samples that are too young to be relevant?

I'm certainly not suggesting that, in hindsight, the said paper now looks fundamentally flawed. In fact, I'd say that it has aged remarkably well, especially considering how fast things are moving in the field of ancient genomics.

But those loose ends really need tying up, one way or another. It's now time.

So someone out there, please, let us know finally if you have the relevant Yamnaya samples. And if you don't, that's OK too, just tell us what you do have. Indeed, it'd be nice know a few basic details about the thousands of samples that have been successfully sequenced in various labs and are waiting to be published. A lot of people would appreciate it.

See also...

Corded Ware as an offshoot of Hungarian Yamnaya (Anthony 2017)

Hungarian Yamnaya > Bell Beakers?

Late PIE ground zero now obvious; location of PIE homeland still uncertain, but...

Saturday, February 23, 2019

Catacomb > Armenia_MLBA


It's now clear, thanks to ancient DNA, that Transcaucasia and surrounds were affected by multiple, and at times significant, population movements from Eastern Europe during the Chalcolithic and Bronze Age periods. Based on the ancient samples from what is now Armenia, I'd say that this process peaked during the Middle Bronze Age. But who exactly were the people who perhaps swarmed south of the Caucasus at this time?

The most likely suspects are the various groups that occupied the southernmost parts of the Pontic-Caspian steppe throughout the Bronze Age. They were associated with the so called Catacomb, Kubano-Tersk and Yamnaya archeological cultures. Below is a Principal Component Analysis (PCA) that compares samples from these cultures with those from Middle to Late Bronze Age Armenia (labeled Armenia_MLBA). The relevant datasheet is available here.


Note that Armenia_MLBA forms a cline that appears to be stretching out towards the Catacomb, Kubano-Tersk, Yamnaya and other Bronze Age steppe groups, and this suggests that it harbors significant and probably recent steppe-related ancestry. But PCA plots based on just two dimensions of genetic variation can be misleading at times, so let's check this out with some formal mixture models using qpAdm.

Armenia_MLBA
Catacomb 0.234±0.028
Kura-Araxes_Kaps 0.766±0.028
chisq 10.723
tail prob 0.826248
Full output

Armenia_MLBA
Kubano-Tersk 0.254±0.030
Kura-Araxes_Kaps 0.746±0.030
chisq 13.535
tail prob 0.633284
Full output

Armenia_MLBA
Kura-Araxes_Kaps 0.768±0.028
Yamnaya_Kalmykia 0.232±0.028
chisq 14.454
tail prob 0.564954
Full output

Armenia_MLBA
Kura-Araxes_Kaps 0.762±0.029
Yamnaya_Caucasus 0.238±0.029
chisq 15.916
tail prob 0.458816
Full output

All of these models are statistically very sound, and even though I ranked the results by "tail prob", there's nothing in the output that clearly points to any one of the southern steppe groups as the obvious source of the steppe-related ancestry in Armenia_MLBA. But, interestingly, Catacomb tops the ranking, and it probably also makes the most sense based simply on Carbon-14 chronology. So, for now, I'm going with Catacomb.

I didn't get a chance yet to investigate this issue in detail with the Global25. Does it contradict the results from my PCA and qpAdm analyses? If anyone reading this would like to take a close look that'd be great. Feel free to post your findings in the comments below. And if the answer is indeed Catacomb, then what language did these Catacomb-derived migrants, or perhaps invaders, speak? If not proto-Armenian then what?

By the way, please be aware that the Kubano-Tersk samples in my analyses are the same individuals as those featured in Wang et al. 2019 under the label "North Caucasus".

See also...

Early chariot drivers of Transcaucasia came from...

Likely Yamnaya incursion(s) into Northwestern Iran

Late PIE ground zero now obvious; location of PIE homeland still uncertain, but...

Saturday, February 9, 2019

Blast from the past: Matters of basic geography


I'm re-posting this article from 2017 for the benefit of some Science News journalists, who are apparently having major problems dealing with basic geography. That's because they think that the Yamnaya culture was located in Asia rather than Eastern Europe. Take my advice and don't read Science News whatever you do. It might rot your brain.

...


The steppe north of the Black Sea in Ukraine has basically always been considered a part of Europe, and just over 100 years ago some guy with a map decided that the steppe between the eastern coast of the Black Sea in Russia and the Ural River in western Kazakhstan should also be Europe.

So nowadays, right or wrong, it's generally accepted that the entire steppe region west of the Ural River, known as the Pontic-Caspian steppe, is in Eastern Europe. Here's a map courtesy of Wikipedia showing how the official boundary between Eastern Europe and Asia has shifted since the 18th century.


But this decision wasn't entirely arbitrary, because the current boundary between Eastern Europe and Asia by and large follows several major geographic barriers, including the Caucasus Mountains, the Caspian Sea and the Ural Mountains. It'd be hard to argue that these barriers haven't had a profound impact across the ages on the character of Europe and its people, and this has probably been known for well over a couple hundred years.


For instance, if we're to trust the most common interpretations of the works of ancient geographers like Hecataeus and Herodotus, then their worlds in some important ways resembled the typical Principal Component Analysis (PCA) of West Eurasian genetic variation. And it seems that they had a pretty good idea where both the strong continental boundaries and fuzzy areas were located.

Below, on the geographic map inspired by Herodotus, Europa or Europe is delineated from much of Asia by the Black Sea, the Caucasus Mountains and the Caspian Sea, while on the genetic map, most European and Asian populations form two, more or less parallel, clusters fairly cleanly separated by empty space (this was first noted in Lazaridis et al. 2013). Indeed, this empty space is the work of the Black Sea, the Caucasus Mountains and the Caspian Sea acting as rather effective barriers to gene flow between Eastern Europe and Asia (see Yunusbayev et al. 2012).


However, on the genetic map, the Iranic Scythians of the Asian steppes straddle my somewhat arbitrary red line separating Europa and Asia, and this is echoed on the Herodotus map by Iranic and related peoples like the Massagetae and Issedones, who inhabit the seemingly undefined part of the world between Europa and Asia east of the Caspian Sea (Mare Caspium).

Nothing really ground breaking, but pretty cool stuff.

On a related note, I've seen the term "mainland Europe" used recently in at least one of the big ancient DNA papers to describe the part of Europe west of the Pontic-Caspian steppe. It seems that the authors wanted to underline the fairly stark genetic difference that existed between most of Europe and the steppe just prior to the expansion of Yamnaya and related steppe herder groups that initiated the formation of the present-day European gene pool.

I can see why they did this, but to my mind they got things backwards. That's because the term mainland implies the opposite of island and/or peninsula, and of course the part of Europe west of the Pontic-Caspian steppe is a relatively narrow strip of land surrounded by water, so it's a peninsula. Let's visualize these two models on a map of Europe courtesy of Wikipedia:


I understand that my model might result in heart palpitations for some readers, especially those from Western Europe, who generally see their part of Europe as core Europe, but I feel that it makes good sense from a purely geographic POV.

See also...

Max Planck scientists: on a mission against geography

Genetic borders are usually linguistic borders too

Monday, June 26, 2017

Matters of geography


The steppe north of the Black Sea in Ukraine has basically always been considered a part of Europe, and just over 100 years ago some guy with a map decided that the steppe between the eastern coast of the Black Sea in Russia and the Ural River in western Kazakhstan should also be Europe.

So nowadays, right or wrong, it's generally accepted that the entire steppe region west of the Ural River, known as the Pontic-Caspian steppe, is in Eastern Europe. Here's a map courtesy of Wikipedia showing how the official boundary between Eastern Europe and Asia has shifted since the 18th century.


But this decision wasn't entirely arbitrary, because the current boundary between Eastern Europe and Asia by and large follows several major geographic barriers, including the Caucasus Mountains, the Caspian Sea and the Ural Mountains. It'd be hard to argue that these barriers haven't had a profound impact across the ages on the character of Europe and its people, and this has probably been known for well over a couple hundred years.


For instance, if we're to trust the most common interpretations of the works of ancient geographers like Hecataeus and Herodotus, then their worlds in some important ways resembled the typical Principal Component Analysis (PCA) of West Eurasian genetic variation. And it seems that they had a pretty good idea where both the strong continental boundaries and fuzzy areas were located.

Below, on the geographic map inspired by Herodotus, Europa or Europe is delineated from much of Asia by the Black Sea, the Caucasus Mountains and the Caspian Sea, while on the genetic map, most European and Asian populations form two, more or less parallel, clusters fairly cleanly separated by empty space (this was first noted in Lazaridis et al. 2013). Indeed, this empty space is the work of the Black Sea, the Caucasus Mountains and the Caspian Sea acting as rather effective barriers to gene flow between Eastern Europe and Asia (see Yunusbayev et al. 2012).


However, on the genetic map, the Iranic Scythians of the Asian steppes straddle my somewhat arbitrary red line separating Europa and Asia, and this is echoed on the Herodotus map by Iranic and related peoples like the Massagetae and Issedones, who inhabit the seemingly undefined part of the world between Europa and Asia east of the Caspian Sea (Mare Caspium).

Nothing really ground breaking, but pretty cool stuff.

On a related note, I've seen the term "mainland Europe" used recently in at least one of the big ancient DNA papers to describe the part of Europe west of the Pontic-Caspian steppe. It seems that the authors wanted to underline the fairly stark genetic difference that existed between most of Europe and the steppe just prior to the expansion of Yamnaya and related steppe herder groups that initiated the formation of the present-day European gene pool.

I can see why they did this, but to my mind they got things backwards. That's because the term mainland implies the opposite of island and/or peninsula, and of course the part of Europe west of the Pontic-Caspian steppe is a relatively narrow strip of land surrounded by water, so it's a peninsula. Let's visualize these two models on a map of Europe courtesy of Wikipedia:


I understand that my model might result in heart palpitations for some readers, especially those from Western Europe, who generally see their part of Europe as core Europe, but I feel that it makes good sense from a purely geographic POV.

See also...

Max Planck scientists: on a mission against geography

Genetic borders are usually linguistic borders too